anu·cle·o·late | \ (ˈ)ā-n(y)ü-ˈklē-ə-ˌlāt 
, -lət, -ˈn(y)ü-klē-ə- \
, -lət, -ˈn(y)ü-klē-ə- \Definition of anucleolate
: lacking a cell nucleolus These findings were corroborated by the anucleolate mutant, which served as a control, since it lacks ribosomal RNA genes.— Walter J. Gehring, Master Control Genes in Development and Evolution … , 1998
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The first known use of anucleolate was in 1955
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