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Throughout the marshes and along sandy beaches, visitors will see pelicans wading around piers with their gular pouches full of fish.—
Amanda Ogle,
Travel + Leisure,
15 Dec. 2025 In both groups, anoles performed a throat-pumping action called gular pumping, which many types of lizards use to supplement their lungs with oxygen.—
Mindy Weisberger,
CNN,
17 Sep. 2024 Herons, nighthawks, pelicans, doves, and owls cool themselves with gular fluttering, a frequent vibration of their throat membranes, which increases airflow and thus increases evaporation.—
Jeff Goodell,
Time,
6 July 2023 The dolphin reorients the fish while still pulling the lip down and expanding the gular area apparently eliciting intraoral pressure reduction, yet the fish almost escapes.—
Jennifer Ouellette,
Ars Technica,
1 Jan. 2023 The way pelicans cool off, Naish and Stiteler said, is by holding the beak open while fluttering the gular pouch with each breath, somewhat similar to how dogs pant – but more efficiently and without loss of body fluids.—
Miriam Fauzia,
USA TODAY,
1 Sep. 2021 Frigate birds soar above us, their balloon-like red gular pouches inflated below their beaks.—
Mary Holland,
Condé Nast Traveler,
30 Aug. 2019