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                                                      : the wedge-shaped piece at the crown of an arch that locks the other pieces in place   see arch illustration                                      
                
                    
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                                                      : something on which associated things depend for support                                      
              
                             
determination, a keystone of the puritan ethic— L. S. Lewis
L. S. Lewis
                         
                
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 or keystone species                                  
              
          
                                                      : a species of plant or animal that produces a major impact (as by predation) on its ecosystem and is considered essential to maintaining optimum ecosystem function or structure                                      
                
                    
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  Merriam-Webster unabridged




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