Definition of keystone
1 : the wedge-shaped piece at the crown of an arch that locks the other pieces in place — see arch illustration
2 : something on which associated things depend for support determination, a keystone of the puritan ethic— L. S. Lewis
3 or keystone species : a species of plant or animal that produces a major impact (as by predation) on its ecosystem and is considered essential to maintaining optimum ecosystem function or structure
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